Revision of the leafy liverwort genus Radula (Porellales, Jungermanniopsida) in Baltic and Bitterfeld amber

The phylogenetically isolated leafy liverwort genus Radula comprises similar to 200 extant species and has a near worldwide distribution. Most species are obligate or facultative epiphytes on tree bark or leaves in humid, tropical or warm-temperate regions. The fossil record of the genus is limited to a few specimens in amber that all deserve re-examination. Based on a revision of Baltic and Bitterfeld amber inclusions from the Geoscientific Museum Gottingen, the Museum fur Naturkunde Berlin, and the collection Gran we accept three fossil Radula species, R. baltica sp. nov., R. obloneolia and R. sphaerocarpoides. Radula baltica is known from only a single piece of Baltic amber, whereas the other two species have been reported in Baltic and Bitterfeld amber. A neotype is designated for Radula oblongifolia. Radula obloneolia and R. sphaerocarpoides are both morphologically variable, yet we are currently unable to discriminate accessions from the different deposits. (C) 2016 Elsevier B.V. All rights reserved.